Evolutionary Neuroscience, Second Edition, is a collection of chapters on brain evolution that combines selected topics from the recent comprehensive reference, Evolution of Nervous Systems (Elsevier, Academic Press, 2017, 9780128040423). The selected chapters cover a broad range of topics, from historical theory, to the most recent deductions from comparative studies of brains. The articles are organized in sections focused on history, concepts and theory, the evolution of brains from early vertebrates to present-day fishes, amphibians, reptiles and birds, the evolution of mammalian brains, and the evolution of primate brains, including human brains. Each chapter is written by a leader or leaders in the field. Specific topics include brain character reconstruction, principles of brain scaling, basic features of vertebrate brains, the evolution of the major sensory systems, other parts of brains, what we can learn from fossils, the origin of neocortex, and the evolution of specializations of human brains. The collection of articles will be interesting to anyone who is curious about how brains evolved from the simpler nervous systems of the first vertebrates into the many different complex forms now found in present-day vertebrates.